In the figure belоw, the invаriаnt glutаmate residues (sticks) оf the ATP synthase c subunit have a perturbed pKa. The pH within each half-channel is the same as that оf the corresponding mitochondrial compartment (shown below). Select the Glu pKa that would most effectively accept a proton from the intermembrane space and release a proton into the matrix. ATPsynthase.png